In the game's main Story Mode, the player controls Gear Head through five different worlds of seven missions each. The Professor finds that his robot Gear Head isn't affected by the hypnosis, so he modifies him to become a top racer, able to beat MetaCog and his gang bosses and recapture the stolen parts and cars. The evil MetaCog attacks Hot Wheels HQ, putting the engineers to sleep with his hypnotic headlight, stealing five cars along with five parts of a new prototype. It shares the general concept and some characters, but has its own story, modes and gameplay style. Hot Wheels Micro Racers features mini Hot Wheels vehicles as the game's playable and computer-controlled racers, with gameplay consisting of arcade-style racing.Released in conjunction with a version on console and PC, Hot Wheels: Velocity X on Game Boy Advance is not an adaptation, but an entirely unique game. The game features three race tracks, each one with their own unique hazards and obstacles: an office, a child's bedroom, and a beach. Three gameplay modes are featured: Single, Championship, and Pursuit. In Single mode, the player can choose any track on which to race.

In Championship, the player must finish each race in first place in order to advance to the next race. Pursuit consists of a single race with three laps the racer in last place explodes at the end of each lap, with the final remaining car being the winner. The player begins with two vehicles, while five more can be unlocked by completing championship races, and by finding secret areas and hidden stars in each track. Each vehicle is based on a real Hot Wheels toy, and each one has different handling characteristics, such as top speed and acceleration. A turbo boost can be used by the player to temporarily speed up, but must recharge after each use.

The game includes a two-player option, in which both players play on a split screen. Scott Steinberg of IGN criticized the game's limited number of race tracks and wrote: 'Tracks, designed to look gigantic in relation to your puny Hot Wheels racer, are neither complex nor lengthy'. Steinberg praised the graphics but wrote that the game 'would be considered less than half of a completed game under normal circumstances.

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